Freshwater microbial ecology with emphasis on primary and secondary productivity of algae and bacteria, nutrient cycling, and interactions of microbiota with other trophic levels in lakes, streams, thermal springs, river floodplains, and wetlands.
Freshwater microbial ecology and nutrient cycling in streams/rivers, lakes, and wetlands; Nitrogen cycle processes with emphasis on nitrate transport and nitrogen fixation; Primary productivity of algae and aquatic vascular plants, secondary productivity of bacteria; Interactions of microbiota with other trophic levels.