Research interests:

  • Mammalian paleontology
  • Quantitative evolution
  • Biotic responses to changing earth systems


  • Vertebrate Paleontology, Paleobiology, Geology, Evolution, Morphometrics

A vertebrate paleontologist, he studies the evolution of mammals and other vertebrates in the fossil record, including trait-based studies of community response to environmental change, geometric morphometric analysis of evolution and morphology, phylogenetics, biogeography, and speciation on regional and continental geographic scales.

Research: Biology; Climate change; Ecology; Evolution; Paleontology; Vertebrates

Research Area:

  • Environmental Science

Research Interests: vertebrate paleontology; comparative morphological variation at the level of the metapopulation; and the effects of Quaternary climate change on vertebrate faunas

Past Affiliations

Lecturer, School of Biological Sciences, Queen Mary, University of London (past)

Lecturer, Molecular & Cellular Biology, Biomedical Sciences, St. Bartholomew's and the Royal London School of Medicine and Dentistry (past)

Fellow, Michigan Society of Fellows, Geological Sciences, University of Michigan (past)

Biological Science, Animal Science, Geology, Geology, Anthropology
PhD, University of California, Berkeley, Paleontology, 1993
BA, University of Texas at Austin, 1987
biological sciences mammalogy earth sciences paleontology ecosystems

Fellow, American Association for the Advancement of Science, 2021-Present

Society of Vertebrate Paleontology
Society for the Study of Evolution
Society of American Naturalists
Society of Systematic Biologists
Palaeontological Association
Paleontological Society