- Mammalian paleontology
- Quantitative evolution
- Biotic responses to changing earth systems
- Vertebrate Paleontology, Paleobiology, Geology, Evolution, Morphometrics
A vertebrate paleontologist, he studies the evolution of mammals and other vertebrates in the fossil record, including trait-based studies of community response to environmental change, geometric morphometric analysis of evolution and morphology, phylogenetics, biogeography, and speciation on regional and continental geographic scales.
Research: Biology; Climate change; Ecology; Evolution; Paleontology; Vertebrates
Research Interests: vertebrate paleontology; comparative morphological variation at the level of the metapopulation; and the effects of Quaternary climate change on vertebrate faunas