Research Interests: Animal ecological physiology; energetic costs of reproduction and embryonic development; sexual dimorphism; herpetology.

Most of my recent work has focused on internal anatomical and physiological dimorphisms related to reproductive cost in spring breeding frogs and salamanders.  I am also interested in  the influence of environmental quality within amniote eggs on hatchling phenotype (turtles and birds), water balance and desiccation tolerance in crayfish, correlates between morphology and interspecific competition in crayfish, the impact of trematode parasitism on fecundity and stored energy in amphibians, and the effects of thermal pollution on native mussel species.



Professor, School of Sciences, Indiana University Kokomo

Assistant Professor, Biology, Indiana University Kokomo
1998 - current

Associate Professor, Natural, Information, and Mathematical Sciences, Indiana University Kokomo
2004 - current

Past Affiliations

Associate Professor, School of Sciences, Indiana University Kokomo

Associate Professor, Master of Arts in Liberal Studies, School of Humanities and Social Sciences, Indiana University Kokomo

Biological Science, Natural Science, Information Science/Systems, Mathematics
PhD, Miami University, Zoology, 1998
MS, Miami University, Zoology, 1995
BA, Kalamazoo College, Biology, Religion, 1991
metabolism biological sciences zoology evolutionary biology thermoregulation reproductive biology biological adaptation energetics terrestrial ecology
American Society of Ichthyologists and Herpetologists
Herpetologists' League
Indiana Academy of Science
Sigma Xi, The Scientific Research Society
Society for Integrative and Comparative Biology
Society for the study of Amphibians and Reptiles