Our research focused on mechanisms of evolution at the gene, genomic, cellular, and phenotypic levels, with special attention being given to the roles of mutation, random genetic drift, and recombination.


  • The Evolution of Genome Architecture
  • The Role of Mutation in Evolution
  • The Role of Recombination in Evolution
  • Methodology for the Analysis of High-Throughput Genomic Data
  • Evolutionary Cell Biology
Molecular Biology, Biological Science
PhD, University of Minnesota, Ecology and Behavioral Biology, 1977
BS, St. Bonaventure University, Biology, 1973