Research interests include both fundamental and applied research in the areas of sensory physiology and pathophysiology. Further research is ongoing into various aspects of animal behavior, including training methods for dogs and behavioral problems of domestic species. More specifically, fairly large-scale studies on factors influencing performance of dogs and dog-handler teams are being done, along with studies of the electrical activity of the olfactory and trigeminal systems and the influence of disease and environmental conditions upon the senses of smell, vision, and taste.