Expertise

My research involves plant anatomy, in particular wood structure. In am interested in the construction and phylogeny of torus-bearing pit membranes. My teaching interests involve plant anatomy and general biology for majors. One area of research in my laboratory involves wood anatomy, and we have a continuing project involving the structure, development, chemical composition and function of pit membranes. A second area of research involves diatoms, their structure and ecology. A final area of interest is biology education for the visually impaired.

Degrees
PhD, University of Wisconsin, Botany, 1976
MS, Ohio State University, Botany, 1972
BS, Ohio State University, Botany, 1969
Keywords
wildlife wood or wood products biological sciences anatomy botany plant physiology or morphology zoology